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Q&A 9. How does an electric eel shock anything in an environment where everything is grounded? The conditions are so different from the usual electrical accident situation where, for example, one stands in a bathtub full of water, which is grounded through the piping, and touches a live wire. I can see how the shock passes through the body in that case, but I do not understand how the same thing happens under water. Wilmington, Delaware
When a voltage is set up between two electrodes in water, most of the current that results will flow in a straight line between the two electrodes. There will, however, be other lines of current flow that loop out into the water surrounding the electrodes. This may be understood by visualizing an electric circuit consisting of many resistors in parallel, some higher in resistance than others. There will be a certain amount of current flowing in the high resistors in spite of the fact that there are low resistance paths available. For a similar, reason current will flow in a three-dimensional pattern surrounding the electrodes in the water even though the path of lowest resistance is directly between the electrodes. If there is organic material near the electrodes it may or may not form a high resistance path with respect to the water around it (pure fresh water is not a good conductor). In any case, if the voltage between the electrodes is high enough, a shock will be registered. The electric eel acts like two electrodes with a battery voltage supply. It is somewhat misleading to think of this situation in terms of the water being part of a special electrical entity called a "ground." It is simply a conductor of a different form from the wires with which we are used to dealing.
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Q&A 10. How do electric fishes produce their electricity?
Carol Stream, Illinois
Fishes that are capable of emitting electric discharges have electric organs composed of multiple stacks of coin-shaped cells called electroplaques. The electroplaques are derived from neuromuscular tissue. Nervous stimulation of each of them produces a small electrical flow, from one face of the coin- shaped cell toward the other. Many electroplaques arranged in a column, all oriented with the innervated side up, will produce a strong net current when stimulated simultaneously. The electroplaque columns are analogous to batteries aligned in series: the more electroplaques, or batteries, in a line, the greater the voltage produced. (Up to 550 volts can be discharged by the South American electric eel, Electrophorus electricus). Also, the more stacks aligned in parallel, the greater the amperage. The ability of electroplaques to generate an electric current is due to the structure of their cell membranes and the activity of chemical pumps. Sodium ions (Na+) are actively pumped out of the cell, creating an electrochemical gradient across the cell membrane. When the resting permeability of the cell membrane is altered (initially by nervous stimulation of the electroplaque surface), there is a net flow inward of the positive sodium ions. The flow of the positive ions toward the negative interior sets up a small electric current. Interesting facts about electric capabilities in fishes are given in "Fishes with Electric Know-How," Sea Frontiers, vol. 21, no. 3, May-June 1975 and in "Living Power Plants," Sea Frontiers, vol. 3, no. 2, June 1957.


출처 : http://helium.vancouver.wsu.edu/~ingalls/eels/index.html
Features
The electric eel seems to have adapted and evolved to its preferred environment. It is able to swim forward and backward with its undulating anal fin. This allows it a better sense of its surroundings. The muddy waters it inhabits contain extremely low oxygen levels.
The fish frequently surfaces and ‘gulps’ 80% of its oxygen through its practically toothless, yet heavily vasculated mouth. The gills filter the remaining 20% of its oxygen intake. An electric eel will die if left in water for more than 20 minutes.
The cloudy water also does not create visual obstacles for the fish either. Not only is the electric eel nocturnal, a fully grown fish can hardly use their beady eyes. Although the youth can see, the adults become increasingly blind due to the constant exposure to the generated electrical field. In fact, the world of the electric eel is entirely electrical.

By producing an electrical current with low-voltage, the electric eel creates an electrical field, which surrounds their elongated bodies. This enables it to navigate and communicate. Electric fish like the electric eel are generally divided into two categories: Strongly electric fish and Weakly electric fish. Strongly electric fish are distinguished by a pulsated-like discharge, as opposed to the Weakly electric fish, with a wavelike discharge.
These discharges are called electric organ discharge (EOD). Strongly electric fish use electric organ discharge to stun prey in addition to navigation, object detection (electrolocation) and communication (electrocommunication.)
Of the Strongly electric fish, the electric eel is set apart. No other electric animal compares to the electric eel’s unique ability to generate such an enormous amount of electricity for a predatory and territorial weapon. A fully grown eel can produce a potential difference of 600 volts (five times an electrical outlet). In contrast, another Strongly electric fish, the torpedo ray, can only discharge approximately half the voltage of the Electrophorus Electricus.
When the electric eel shocks its prey, it either stuns and paralyzes it or kills it. This would be evolutionarily beneficial to the toothless fish so that the prey does not present a struggle to its predator. The electric eel also shocks itself in the process, yet tolerates the current. Its place in the electrical current flow decreases the effects of the shock, as does its thick, course skin.
This extraordinary ability has naturally drawn a great deal of attention as to how electrical generation of this magnitude occurs within a single animal. Here we’ll explore the underlying mechanisms of the electrical discharge that set electric eels apart from all other animals.
Electric Organs and Electrocytes
The vital organs of the electric eel are located entirely in the first 1/5 of its body, leaving the remaining 4/5 tail occupied by three electric organs.
The Sachs’ organ is where low-voltage pulses are emitted for electrolocation and navigation.
The Hunter and Main organs are where the high-voltage emission occurs.
Electric organs are made up of cells called Electrocytes. Some scientists believe these cells are derivative of a muscle-cell since nerve cells synapse onto them and they behave much like a muscle-cell post-synaptically. However, they are unlike muscle cells in that they don’t contract. Flat and disk-like, the electrocytes are stacked in a sequence much like a dry-cell battery, with the head as the positive pole and the tail as the negative pole. Each electrocyte generates .15 volts, which is a very small amount. However, when 4,000 electrocytes are lined up generating electricity at the exact same time at .15 volts each, the shock equals at least 600 volts, which can paralyze or kill a human, especially after repeated shocks.

At rest, the Na+K+ pump, concentration and electrical gradient keeps the inside of the electrocyte at a resting potential of .08 volt. When the electric eel electrolocates its prey, the brain sends a signal through the nervous system to the electric organs. Acetylcholine is dropped onto the electrocyte which binds to the corresponding receptor on the ion. This opens the ion channels of the cell, allowing Na+ to rush in. The cell then depolarizes, momentarily reversing the charge, and fires.
Since the electrocytes are lined up, current flows through like a battery, emitting a charge. However, they must all discharge at the same time. The electric eel’s design resolves this at the neuronal connections by delaying the signals and action potentials. The closer connections have longer and thinner pathways, which decelerates the signal and allows all electrocytes to synchronize their discharge. 

